2004-present;Principal Scientist within the Disease Control division of the Moredun Research Institute (MRI). 2002-2003; Project Leader/PI, Dept of Veterinary Pathology, University of Edinburgh. Wellcome Trust funded project focusing on novel class II MHC genes in ruminants.2001-2002; Scientific Adviser, Agriculture and Biological Research Group (Scottish Government). 1994-2000 Associate Scientist in immunology and vaccine development, International Livestock Research Institute (ILRI), Nairobi, Kenya; Working on the genetic control of the response to challenge following vaccination with a subunit vaccine targeting the protozoan parasite of cattle Theileria parva. 1997-2004 PhD student (University of Edinburgh) and Postdoctoral Scientist (MRI) working on Characterisation of the class II MHC genes in sheep.